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ENZYME

ENZYME entry: EC 3.4.19.12

Accepted Name
ubiquitinyl hydrolase 1
Alternative Name(s)
ubiquitin C-terminal hydrolase
ubiquitin thiolesterase
Reaction catalysed
Thiol-dependent hydrolysis of ester, thioester, amide, peptide and isopeptide bonds formed by the C-terminal Gly of ubiquitin (a 76-residue protein attached to proteins as an intracellular targeting signal)
Comment(s)
  • Links to polypeptides smaller than 60 residues are hydrolyzed more readily than those to larger polypeptides.
  • Isoforms exist with quantitatively different specificities among the best known being UCH-L1 and UCH-L3, major proteins of the brain of mammals.
  • Inhibited by ubiquitin aldehyde (in which Gly76 is replaced by aminoacetaldehyde).
  • Belongs to peptidase family C12.
Cross-references
BRENDA3.4.19.12
EC2PDB3.4.19.12
ExplorEnz3.4.19.12
PRIAM enzyme-specific profiles3.4.19.12
KEGG Ligand Database for Enzyme Nomenclature3.4.19.12
IUBMB Enzyme Nomenclature3.4.19.12
IntEnz3.4.19.12
MEDLINEFind literature relating to 3.4.19.12
MetaCyc3.4.19.12
Rhea expert-curated reactions3.4.19.12
UniProtKB/Swiss-Prot
Q9M391, ATX3H_ARATHQ9H3M9, ATX3L_HUMANQ60XN1, ATX3_CAEBR
O17850, ATX3_CAEELQ9W689, ATX3_CHICKP54252, ATX3_HUMAN
Q9CVD2, ATX3_MOUSEQ8LQ36, ATX3_ORYSJO35815, ATX3_RAT
A2VDM8, BAP1_BOVINC4A0D9, BAP1_BRAFLQ5F3N6, BAP1_CHICK
A1L2G3, BAP1_DANREQ92560, BAP1_HUMANQ99PU7, BAP1_MOUSE
D3ZHS6, BAP1_RATQ52L14, BAP1_XENLAQ66JB6, BAP1_XENTR
Q17N72, CALYP_AEDAEB0W2R4, CALYP_CULQUB3MIV9, CALYP_DROAN
B3NPV7, CALYP_DROERB4JW98, CALYP_DROGRQ7K5N4, CALYP_DROME
B4KT51, CALYP_DROMOB4GAM2, CALYP_DROPEQ291J4, CALYP_DROPS
B4HST0, CALYP_DROSEB4QHH0, CALYP_DROSIB4LQ24, CALYP_DROVI
B4P6P6, CALYP_DROYAA1CIL1, CREB_ASPCLB0Y4P5, CREB_ASPFC
B8NSV5, CREB_ASPFNQ4WQI1, CREB_ASPFUA2Q9N1, CREB_ASPNC
Q2UUG8, CREB_ASPORQ0CT11, CREB_ASPTNQ96V54, CREB_EMENI
A1CW53, CREB_NEOFIQ1RMU2, CYLD_BOVINQ9NQC7, CYLD_HUMAN
Q80TQ2, CYLD_MOUSEQ5RED8, CYLD_PONABQ66H62, CYLD_RAT
Q5ZIV7, DESI2_CHICKQ6DC39, DESI2_DANREQ9BSY9, DESI2_HUMAN
Q9D291, DESI2_MOUSEA3QRX8, DESI2_PIGQ5R456, DESI2_PONAB
Q5XIT6, DESI2_RATQ5PQ09, DESI2_XENLAO00303, EIF3F_HUMAN
Q4R5B8, EIF3F_MACFAQ9DCH4, EIF3F_MOUSEA5A6I3, EIF3F_PANTR
P55824, FAF_DROMEQ5EAE5, JOS1_BOVINQ15040, JOS1_HUMAN
Q9DBJ6, JOS1_MOUSEQ5R739, JOS1_PONABQ5BJY4, JOS1_RAT
Q8TAC2, JOS2_HUMANQ9CR30, JOS2_MOUSEO82391, JOSL_ARATH
Q9W422, JOSL_DROMEO36414, LTP_ALHV1Q1HVH9, LTP_EBVA8
P03186, LTP_EBVB9Q3KSU8, LTP_EBVGP28955, LTP_EHV1B
Q66666, LTP_EHV2Q9E6N3, LTP_GAHVMP16785, LTP_HCMVA
Q6SW84, LTP_HCMVMD5LX59, LTP_HCMVTP10220, LTP_HHV11
P89459, LTP_HHV2HP30002, LTP_HHV6GP52340, LTP_HHV6U
Q9QJ37, LTP_HHV6ZP52362, LTP_HHV7JQ2HR64, LTP_HHV8P
O41965, LTP_MHV68A8E1C4, LTP_MUHVKQ6UDJ5, LTP_PSHV1
Q01056, LTP_SHV21P09278, LTP_VZVDQ4JQX9, LTP_VZVO
Q6TQR6, L_CCHFIQ66431, L_DUGBAQ2KJ22, MINY1_BOVIN
A3KQS4, MINY1_DANREQ8N5J2, MINY1_HUMANQ76LS9, MINY1_MOUSE
Q5R7G8, MINY1_PONABQ5BJQ2, MINY1_RATQ2KI23, MINY2_BOVIN
Q8NBR6, MINY2_HUMANQ6PDI6, MINY2_MOUSEQ0IIH8, MINY3_BOVIN
A0AUR5, MINY3_DANREQ9VWN5, MINY3_DROMEQ9H8M7, MINY3_HUMAN
Q4R528, MINY3_MACFAQ9CV28, MINY3_MOUSEQ6NX27, MINY3_XENTR
A1A4L4, MINY4_BOVINQ4G0A6, MINY4_HUMANQ3UQI9, MINY4_MOUSE
Q5RF72, MINY4_PONABQ0VA42, MINY4_XENTRO13979, MU105_SCHPO
Q9VVR1, NOT_DROMEQ9LZF7, OTU10_ARATHQ0V869, OTU11_ARATH
Q9SGA5, OTU12_ARATHQ8LG98, OTU1_ARATHQ05B57, OTU1_BOVIN
Q5F3A6, OTU1_CHICKQ567B1, OTU1_DANREQ55BI3, OTU1_DICDI
Q9VRJ9, OTU1_DROMEQ29FC9, OTU1_DROPSQ5VVQ6, OTU1_HUMAN
Q8CB27, OTU1_MOUSEQ32Q05, OTU1_RATO13974, OTU1_SCHPO
Q0IH43, OTU1_XENLAP43558, OTU1_YEASTQ9LPT6, OTU2_ARATH
Q9UUK3, OTU2_SCHPOQ8GYW0, OTU3_ARATHQ8LBZ4, OTU4_ARATH
Q08BW0, OTU5A_DANREQ7ZX21, OTU5A_XENLAQ640H3, OTU5B_XENLA
Q9LYC7, OTU5_ARATHQ7L8S5, OTU6A_HUMANQ6IE21, OTU6A_MOUSE
Q5ZIP6, OTU6B_CHICKQ7ZV00, OTU6B_DANREQ8N6M0, OTU6B_HUMAN
Q8K2H2, OTU6B_MOUSEQ6GM06, OTU6B_XENLAQ5M8L0, OTU6B_XENTR
Q9XIP2, OTU6_ARATHQ8TE49, OTU7A_HUMANQ8R554, OTU7A_MOUSE
Q6GQQ9, OTU7B_HUMANB2RUR8, OTU7B_MOUSEF4K3M6, OTU7_ARATH
O80949, OTU8_ARATHQ8LBW2, OTU9_ARATHQ96FW1, OTUB1_HUMAN
Q7TQI3, OTUB1_MOUSEB2RYG6, OTUB1_RATQ96DC9, OTUB2_HUMAN
Q9CQX0, OTUB2_MOUSEA8XDJ2, OTUBL_CAEBRQ9XVR6, OTUBL_CAEEL
Q9VL00, OTUBL_DROMEQ5VV17, OTUD1_HUMANQ9CUB6, OTUD1_MOUSE
Q5T2D3, OTUD3_HUMANB1AZ99, OTUD3_MOUSEQ01804, OTUD4_HUMAN
B2RRE7, OTUD4_MOUSEQ96G74, OTUD5_HUMANQ3U2S4, OTUD5_MOUSE
Q2YDU3, OTUD5_RATQ6GL44, OTUD5_XENTRQ96BN8, OTUL_HUMAN
Q3UCV8, OTUL_MOUSEQ155Z9, POLG_SVV1P10358, POLR_TYMV
P20128, POLR_TYMVAP28477, POLR_TYMVCQ9VC56, PUF_DROME
P0C6W1, R1AB_BC133P0C6V9, R1AB_BC279P0C6W0, R1AB_BC512
P0C6W2, R1AB_BCHK3P0C6W3, R1AB_BCHK4P0C6W4, R1AB_BCHK5
P0C6W5, R1AB_BCHK9P0C6W6, R1AB_BCRP3P0C6W7, R1AB_CVBEN
P0C6W8, R1AB_CVBLUP0C6W9, R1AB_CVBMP0C6X0, R1AB_CVBQ
P0C6X1, R1AB_CVH22P0C6X2, R1AB_CVHN1P0C6X3, R1AB_CVHN2
P0C6X4, R1AB_CVHN5P0C6X5, R1AB_CVHNLP0C6X6, R1AB_CVHOC
P0C6X8, R1AB_CVM2P0C6X9, R1AB_CVMA5P0C6Y0, R1AB_CVMJH
P0C6Y5, R1AB_CVPPUQ98VG9, R1AB_FIPVP0C6Y1, R1AB_IBVB
P0C6Y2, R1AB_IBVBCP0C6Y3, R1AB_IBVMK9N7C7, R1AB_MERS1
P0C6Y4, R1AB_PEDV7P0C6X7, R1AB_SARSP0DTD1, R1AB_SARS2
P0C6F7, R1A_BC133P0C6F5, R1A_BC279P0C6F6, R1A_BC512
P0C6F8, R1A_BCHK3P0C6T4, R1A_BCHK4P0C6T5, R1A_BCHK5
P0C6T6, R1A_BCHK9P0C6T7, R1A_BCRP3P0C6T8, R1A_CVBEN
P0C6T9, R1A_CVBLUP0C6U0, R1A_CVBMP0C6U1, R1A_CVBQ
P0C6U2, R1A_CVH22P0C6U3, R1A_CVHN1P0C6U4, R1A_CVHN2
P0C6U5, R1A_CVHN5P0C6U6, R1A_CVHNLP0C6U7, R1A_CVHOC
P0C6U9, R1A_CVM2P0C6V0, R1A_CVMA5P0C6V1, R1A_CVMJH
P0C6V2, R1A_CVPPUP0C6V3, R1A_IBVBP0C6V4, R1A_IBVBC
P0C6V5, R1A_IBVMK9N638, R1A_MERS1P0C6V6, R1A_PEDV7
P0C6U8, R1A_SARSP0DTC1, R1A_SARS2P43588, RPN11_YEAST
P19811, RPOA_EAVBUQ9YN02, RPOA_PRRS1Q8B912, RPOA_PRRSB
Q04561, RPOA_PRRSLQ9WJB2, RPOA_PRRSRA0MD28, RPOA_PRRSS
A6YQT5, RPOA_PRRSVP21580, TNAP3_HUMANQ4R8W3, TNAP3_MACFA
Q60769, TNAP3_MOUSEQ9VH90, TRBID_DROMEQ7RTZ2, U17L1_HUMAN
Q6R6M4, U17L2_HUMANA6NCW0, U17L3_HUMANA8MUK1, U17L5_HUMAN
Q6QN14, U17L6_HUMANC9JJH3, U17LA_HUMANC9JVI0, U17LB_HUMAN
G5E8I7, U17LC_MOUSEC9JLJ4, U17LD_HUMANC9J2P7, U17LF_HUMAN
D6RBQ6, U17LH_HUMAND6R9N7, U17LI_HUMAND6RCP7, U17LJ_HUMAN
D6RJB6, U17LK_HUMAND6R901, U17LL_HUMAND6RA61, U17LM_HUMAN
Q0WX57, U17LO_HUMANQ61068, U17PA_MOUSEE9Q9U0, U17PB_MOUSE
G5E8G2, U17PD_MOUSEQ7M764, U17PE_MOUSEQ2NL57, UB10A_XENLA
Q7ZXR7, UB10B_XENLAQ52KZ6, UB12A_XENLAQ5M981, UB12B_XENLA
Q6GNI6, UB22A_XENLAQ6DCJ1, UB22B_XENLAQ09444, UBH4_CAEEL
P35127, UBL1_YEASTQ10171, UBLH1_SCHPOQ9UUB6, UBLH2_SCHPO
Q9ZSB5, UBP10_ARATHA5PJS6, UBP10_BOVINQ5ZJN4, UBP10_CHICK
Q14694, UBP10_HUMANP52479, UBP10_MOUSEQ3KR59, UBP10_RAT
Q6DIJ4, UBP10_XENTRP53874, UBP10_YEASTQ9MAQ3, UBP11_ARATH
Q01988, UBP11_CANLFP51784, UBP11_HUMANQ99K46, UBP11_MOUSE
Q5D006, UBP11_RATQ9UUD6, UBP11_SCHPOP36026, UBP11_YEAST
Q9FPT1, UBP12_ARATHA5D9H7, UBP12_BOVINA4FUN7, UBP12_DANRE
O75317, UBP12_HUMANQ9D9M2, UBP12_MOUSEC0HB46, UBP12_SALSA
O60079, UBP12_SCHPOP39538, UBP12_YEASTQ84WU2, UBP13_ARATH
E1BMF7, UBP13_BOVINE1BY77, UBP13_CHICKF1QFS9, UBP13_DANRE
Q92995, UBP13_HUMANQ5BKP2, UBP13_MOUSEF6V6I0, UBP13_XENTR
P38187, UBP13_YEASTQ8L6Y1, UBP14_ARATHQ0IIF7, UBP14_BOVIN
Q17361, UBP14_CAEELP54578, UBP14_HUMANQ9JMA1, UBP14_MOUSE
P60051, UBP14_PANTRP40826, UBP14_RABITQ11119, UBP14_SCHPO
P38237, UBP14_YEASTQ9FPS9, UBP15_ARATHQ2HJE4, UBP15_BOVIN
Q9Y4E8, UBP15_HUMANQ8R5H1, UBP15_MOUSEQ0E2F9, UBP15_ORYSJ
Q9R085, UBP15_RATF6Z5C0, UBP15_XENTRP50101, UBP15_YEAST
Q9SB51, UBP16_ARATHQ08DA3, UBP16_BOVINA8HAL1, UBP16_DANRE
P0CAQ1, UBP16_EMENIQ9Y5T5, UBP16_HUMANQ4R6X7, UBP16_MACFA
Q99LG0, UBP16_MOUSEQ2KJ09, UBP16_RATO74442, UBP16_SCHPO
Q6PAW2, UBP16_XENLAQ0VA64, UBP16_XENTRQ02863, UBP16_YEAST
Q9FKP5, UBP17_ARATHQ67XW5, UBP18_ARATHQ9UMW8, UBP18_HUMAN
Q9WTV6, UBP18_MOUSEQ5RE63, UBP18_PONABQ9SJA1, UBP19_ARATH
O94966, UBP19_HUMANQ3UJD6, UBP19_MOUSEQ6J1Y9, UBP19_RAT
Q9FPT5, UBP1_ARATHQ29RP1, UBP1_BOVINO94782, UBP1_HUMAN
Q8BJQ2, UBP1_MOUSEQ569C3, UBP1_RATQ9USM5, UBP1_SCHPO
P25037, UBP1_YEASTQ9FPS7, UBP20_ARATHA7Z056, UBP20_BOVIN
A5PN09, UBP20_DANREQ9Y2K6, UBP20_HUMANQ8C6M1, UBP20_MOUSE
Q5R5Z6, UBP20_PONABB1WBD7, UBP20_XENLAA0JM59, UBP20_XENTR
Q9FIQ1, UBP21_ARATHQ2KJ72, UBP21_BOVINQ9UK80, UBP21_HUMAN
Q9QZL6, UBP21_MOUSEB2GUX4, UBP21_RATQ9LEW0, UBP22_ARATH
P0C8Z3, UBP22_BOVINA6H8I0, UBP22_DANREQ9UPT9, UBP22_HUMAN
Q5DU02, UBP22_MOUSEQ9FPS4, UBP23_ARATHQ9FPS3, UBP24_ARATH
Q9UPU5, UBP24_HUMANB1AY13, UBP24_MOUSEQ9FPS2, UBP25_ARATH
Q9UHP3, UBP25_HUMANP57080, UBP25_MOUSEQ9SCJ9, UBP26_ARATH
Q9BXU7, UBP26_HUMANQ99MX1, UBP26_MOUSEA2XDG4, UBP26_ORYSI
A3AF13, UBP26_ORYSJQ9FPS0, UBP27_ARATHA6NNY8, UBP27_HUMAN
Q8CEG8, UBP27_MOUSEQ5ZID5, UBP28_CHICKQ96RU2, UBP28_HUMAN
Q5I043, UBP28_MOUSED3ZJ96, UBP28_RATQ9HBJ7, UBP29_HUMAN
Q9ES63, UBP29_MOUSEQ8W4N3, UBP2_ARATHQ2KHV7, UBP2_BOVIN
O57429, UBP2_CHICKQ8IQ27, UBP2_DROMEO75604, UBP2_HUMAN
O42726, UBP2_KLULAO88623, UBP2_MOUSEQ5U349, UBP2_RAT
Q9P3U0, UBP2_SCHPOQ01476, UBP2_YEASTA2BGT0, UBP30_DANRE
Q9W462, UBP30_DROMEQ70CQ3, UBP30_HUMANQ3UN04, UBP30_MOUSE
D3ZPG5, UBP30_RATA4QNN3, UBP30_XENTRQ70CQ4, UBP31_HUMAN
M9PD06, UBP32_DROMEQ8NFA0, UBP32_HUMANF8VPZ3, UBP32_MOUSE
A6QNM7, UBP33_BOVINA5PMR2, UBP33_DANREQ8TEY7, UBP33_HUMAN
Q8R5K2, UBP33_MOUSEQ5REG5, UBP33_PONABQ28CN3, UBP33_XENTR
Q70CQ2, UBP34_HUMANQ6ZQ93, UBP34_MOUSEQ9P2H5, UBP35_HUMAN
B3M3M6, UBP36_DROANB3NC86, UBP36_DROERB4IXE0, UBP36_DROGR
Q9VRP5, UBP36_DROMEB4KXJ5, UBP36_DROMOQ2LZB1, UBP36_DROPS
B4HUI5, UBP36_DROSEB4QIS3, UBP36_DROSIB4LG38, UBP36_DROVI
B4MLR8, UBP36_DROWIB4PIW8, UBP36_DROYAQ9P275, UBP36_HUMAN
B1AQJ2, UBP36_MOUSEF1N5V1, UBP37_BOVINE2RK09, UBP37_CANLF
E1C213, UBP37_CHICKE7F6T8, UBP37_DANREQ86T82, UBP37_HUMAN
Q8C0R0, UBP37_MOUSEF1SRY5, UBP37_PIGQ8NB14, UBP38_HUMAN
Q8BW70, UBP38_MOUSEQ53GS9, UBP39_HUMANQ3TIX9, UBP39_MOUSE
Q5R761, UBP39_PONABO24454, UBP3_ARATHQ9Y6I4, UBP3_HUMAN
Q91W36, UBP3_MOUSEO94269, UBP3_SCHPOQ01477, UBP3_YEAST
Q9NVE5, UBP40_HUMANQ8BWR4, UBP40_MOUSEE1B9W9, UBP42_BOVIN
Q9H9J4, UBP42_HUMANB2RQC2, UBP42_MOUSED3ZU96, UBP42_RAT
Q70EL4, UBP43_HUMANQ8BUM9, UBP43_MOUSED2HBJ8, UBP44_AILME
Q7ZUM8, UBP44_DANREQ9H0E7, UBP44_HUMANQ8C2S0, UBP44_MOUSE
Q0V9G5, UBP44_XENTRE9QG68, UBP45_DANREQ70EL2, UBP45_HUMAN
Q8K387, UBP45_MOUSEP34547, UBP46_CAEELA5WWB0, UBP46_DANRE
P62068, UBP46_HUMANP62069, UBP46_MOUSEQ5RBQ4, UBP46_PONAB
F1M625, UBP46_RATE1C1R4, UBP47_CHICKQ96K76, UBP47_HUMAN
Q8BY87, UBP47_MOUSEQ5U252, UBP47_XENLAA1A5G2, UBP47_XENTR
G5ECC7, UBP48_CAEELQ5ZM45, UBP48_CHICKA0A0R4IB93, UBP48_DANRE
Q86UV5, UBP48_HUMANQ3V0C5, UBP48_MOUSEQ76LT8, UBP48_RAT
Q70CQ1, UBP49_HUMANQ6P9L4, UBP49_MOUSEQ8LAM0, UBP4_ARATH
A6QR55, UBP4_BOVINQ6FQF0, UBP4_CANGAQ754R5, UBP4_EREGS
Q13107, UBP4_HUMANQ9HFS7, UBP4_KLULAP35123, UBP4_MOUSE
Q5RCD3, UBP4_PONABB2GUZ1, UBP4_RATO60139, UBP4_SCHPO
A7TGY3, UBP4_VANPOB3LGK1, UBP4_YEAS1A6ZY34, UBP4_YEAS7
P32571, UBP4_YEASTQ70EL3, UBP50_HUMANQ4R6D3, UBP50_MACFA
Q6P8X6, UBP50_MOUSEQ70EK9, UBP51_HUMANB1AY15, UBP51_MOUSE
Q70EL1, UBP54_HUMANQ8BL06, UBP54_MOUSEQ6IE24, UBP54_RAT
O22207, UBP5_ARATHP45974, UBP5_HUMANP56399, UBP5_MOUSE
Q5R407, UBP5_PONABP39944, UBP5_YEASTQ9W4C3, UBP64_DROME
Q949Y0, UBP6_ARATHP35125, UBP6_HUMANQ92353, UBP6_SCHPO
P43593, UBP6_YEASTQ9UTT1, UBP71_SCHPOQ09879, UBP72_SCHPO
Q84WC6, UBP7_ARATHQ60MK8, UBP7_CAEBRQ7JKC3, UBP7_CAEEL
Q6U7I1, UBP7_CHICKQ9VYQ8, UBP7_DROMEQ93009, UBP7_HUMAN
Q6A4J8, UBP7_MOUSEQ4VSI4, UBP7_RATQ9P7S5, UBP7_SCHPO
P40453, UBP7_YEASTQ9C585, UBP8_ARATHP40818, UBP8_HUMAN
Q80U87, UBP8_MOUSEQ09738, UBP8_SCHPOP50102, UBP8_YEAST
Q93Y01, UBP9_ARATHQ9P7V9, UBP9_SCHPOP39967, UBP9_YEAST
P54201, UBPA_DICDIO96612, UBPB_DICDIQ24574, UBPE_DROME
Q8WT44, UBPH_CAEELQ5UQR3, UBPL1_MIMIVQ5UPW7, UBPL2_MIMIV
Q09931, UBPY_CAEELQ9FFF2, UCH1_ARATHO04482, UCH2_ARATH
Q8IIJ6, UCH37_PLAF7Q8GWE1, UCH3_ARATHP23356, UCHL1_BOVIN
Q9GM50, UCHL1_HORSEP09936, UCHL1_HUMANQ60HC8, UCHL1_MACFA
P50103, UCHL1_MONDOQ9R0P9, UCHL1_MOUSEQ6SEG5, UCHL1_PIG
Q00981, UCHL1_RATQ2TBG8, UCHL3_BOVINP15374, UCHL3_HUMAN
Q9JKB1, UCHL3_MOUSEQ06AB3, UCHL3_PIGQ8IKM8, UCHL3_PLAF7
Q91Y78, UCHL3_RATP58321, UCHL4_MOUSEQ9XSJ0, UCHL5_BOVIN
Q54N38, UCHL5_DICDIQ9Y5K5, UCHL5_HUMANQ9WUP7, UCHL5_MOUSE
Q06AT3, UCHL5_PIGO01391, UCHL_APLCAQ54T48, UCHL_DICDI
P35122, UCHL_DROMEC9JPN9, UL17C_HUMANQ6NTR6, UP44A_XENLA
Q5XGZ2, UP44B_XENLAQ93008, USP9X_HUMANP70398, USP9X_MOUSE
O00507, USP9Y_HUMANF8VPU6, USP9Y_MOUSEQ96JH7, VCIP1_HUMAN
Q8CDG3, VCIP1_MOUSEQ8CF97, VCIP1_RATQ196X6, VF232_IIV3
Q91FU0, VF232_IIV6P22856, VL96_IRV1Q08930, YP191_YEAST
A6QP16, ZRAN1_BOVINQ9UGI0, ZRAN1_HUMANQ7M760, ZRAN1_MOUSE
B1H2Q2, ZRAN1_XENTRQ5U595, ZRN1A_XENLAA0JMQ9, ZRN1B_DANRE
Q6NUB7, ZRN1B_XENLAQ3SWY8, ZUP1_BOVINQ96AP4, ZUP1_HUMAN
Q4R4A2, ZUP1_MACFAQ3T9Z9, ZUP1_MOUSEQ5U2S3, ZUP1_RAT

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