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ENZYME

ENZYME entry: EC 4.3.2.10

Accepted Name
imidazole glycerol-phosphate synthase
Alternative Name(s)
IGP synthase
Reaction catalysed
5-[(5-phospho-1-deoxy-D-ribulos-1-ylimino)methylamino]-1-(5-phospho-beta-D-ribosyl)imidazole-4-carboxamide + L-glutamine <=> 5-amino-1-(5-phospho-beta-D-ribosyl)imidazole-4-carboxamide + D-erythro-1-(imidazol-4-yl)glycerol 3-phosphate + H(+) + L-glutamate
Comment(s)
  • The enzyme is involved in histidine biosynthesis, as well as purine nucleotide biosynthesis.
  • The enzymes from archaea and bacteria are heterodimeric.
  • A glutaminase component (cf. EC 3.5.1.2) produces an ammonia molecule that is transferred by a 25 A tunnel to a cyclase component, which adds it to the imidazole ring, leading to lysis of the molecule and cyclization of one of the products.
  • The glutminase subunit is only active within the dimeric complex.
  • In fungi and plants the two subunits are combined into a single polypeptide.
Cross-references
BRENDA4.3.2.10
EC2PDB4.3.2.10
ExplorEnz4.3.2.10
PRIAM enzyme-specific profiles4.3.2.10
KEGG Ligand Database for Enzyme Nomenclature4.3.2.10
IUBMB Enzyme Nomenclature4.3.2.10
IntEnz4.3.2.10
MEDLINEFind literature relating to 4.3.2.10
MetaCyc4.3.2.10
Rhea expert-curated reactions4.3.2.10
UniProtKB/Swiss-Prot
Q9SZ30, HIS4_ARATHQ0P8U2, HIS51_CAMJEA1W0U9, HIS51_CAMJJ
Q5HT90, HIS51_CAMJRQ5X6Z7, HIS51_LEGPAQ5ZXI1, HIS51_LEGPH
Q5WYE9, HIS51_LEGPLP61782, HIS51_METMPQ3SWE9, HIS51_NITWN
Q7U924, HIS51_PARMWQ7V959, HIS51_PROMMQ9HU42, HIS51_PSEAE
Q7MLS3, HIS51_VIBVYQ9PM75, HIS52_CAMJEQ5HSJ1, HIS52_CAMJR
Q5X5X2, HIS52_LEGPAQ5ZW90, HIS52_LEGPHQ5WX94, HIS52_LEGPL
P61783, HIS52_METMPQ3SPZ6, HIS52_NITWNQ7U899, HIS52_PARMW
Q7V6M5, HIS52_PROMMP72138, HIS52_PSEAEQ7MPP4, HIS52_VIBVY
Q7MPK8, HIS53_VIBVYB0BYP8, HIS5_ACAM1Q6F7A7, HIS5_ACIAD
P58788, HIS5_AGRFCQ5E635, HIS5_ALIF1Q5WDI0, HIS5_ALKCK
O66943, HIS5_AQUAEO28019, HIS5_ARCFUQ5P793, HIS5_AROAE
P18785, HIS5_AZOBRC3P504, HIS5_BACAAC3L9P9, HIS5_BACAC
A0RBM0, HIS5_BACAHQ81T60, HIS5_BACANB7JFZ3, HIS5_BACC0
P61778, HIS5_BACC1B7INA1, HIS5_BACC2C1EMQ5, HIS5_BACC3
B7HHG3, HIS5_BACC4B7HKD2, HIS5_BACC7A7GMU8, HIS5_BACCN
B9IUZ8, HIS5_BACCQQ81G04, HIS5_BACCRQ63DX0, HIS5_BACCZ
Q5LBD5, HIS5_BACFNQ64RT0, HIS5_BACFRQ6HLE5, HIS5_BACHK
Q65EG1, HIS5_BACLDA9VLH5, HIS5_BACMKA8FHR1, HIS5_BACP2
O34565, HIS5_BACSUQ8A7Z4, HIS5_BACTNA7Z964, HIS5_BACVZ
P60597, HIS5_BDEBAQ8G4S6, HIS5_BIFLOQ7VQW7, HIS5_BLOFL
Q492K0, HIS5_BLOPBQ7WDY1, HIS5_BORBRQ7W2Y1, HIS5_BORPA
Q7VSY8, HIS5_BORPEQ89WM6, HIS5_BRADUQ2YQZ0, HIS5_BRUA2
Q57AH5, HIS5_BRUABQ8YE35, HIS5_BRUMEQ8FY09, HIS5_BRUSU
P57204, HIS5_BUCAIQ9ZHE3, HIS5_BUCAPP59501, HIS5_BUCBP
Q84I56, HIS5_BUCDNQ84I55, HIS5_BUCMHQ84I54, HIS5_BUCSC
Q39K87, HIS5_BURL3Q845U9, HIS5_BURM1Q62GE3, HIS5_BURMA
Q3JN00, HIS5_BURP1Q63Q90, HIS5_BURPSQ8R883, HIS5_CALS4
Q3AD50, HIS5_CARHZQ9A231, HIS5_CAUVCO33565, HIS5_CERS4
A9WD25, HIS5_CHLAAB8G9S3, HIS5_CHLADQ3AT63, HIS5_CHLCH
Q3B286, HIS5_CHLL3B9LI74, HIS5_CHLSYQ8KF56, HIS5_CHLTE
Q7P0F1, HIS5_CHRVOQ97KI0, HIS5_CLOABQ47XB5, HIS5_COLP3
P60598, HIS5_CORDIQ8FNZ6, HIS5_COREFO69043, HIS5_CORGL
Q4JW55, HIS5_CORJKQ46WL6, HIS5_CUPPJQ6A8L2, HIS5_CUTAK
Q9TLQ8, HIS5_CYACAQ85FY4, HIS5_CYAM1B8HUR1, HIS5_CYAP4
P48262, HIS5_CYAPAQ47AM1, HIS5_DECARQ3Z7F2, HIS5_DEHM1
Q3ZY50, HIS5_DEHMCQ9RX89, HIS5_DEIRAB8J215, HIS5_DESDA
B8FP22, HIS5_DESHDQ24QJ3, HIS5_DESHYP61779, HIS5_DESVH
B8DQX7, HIS5_DESVMA1VGZ0, HIS5_DESVVP58237, HIS5_ECO57
P60596, HIS5_ECOL6P60595, HIS5_ECOLIQ9P4P9, HIS5_EMENI
B1YL11, HIS5_EXIS2Q5KVC8, HIS5_GEOKAQ39YP4, HIS5_GEOMG
P60599, HIS5_GEOSLC5D7P0, HIS5_GEOSWA4ISR4, HIS5_GEOTN
B7KGN3, HIS5_GLOC7Q7NNK4, HIS5_GLOVIQ5FTN5, HIS5_GLUOX
Q0BPX1, HIS5_GRABCQ4QN71, HIS5_HAEI8P44340, HIS5_HAEIN
Q9K6Z4, HIS5_HALH5Q5UYV3, HIS5_HALMAQ9HNI6, HIS5_HALSA
Q7VIW9, HIS5_HELHPA9B5I2, HIS5_HERA2Q31E66, HIS5_HYDCU
Q5QWQ7, HIS5_IDILOB3WED3, HIS5_LACCBQ02132, HIS5_LACLA
Q039B3, HIS5_LACP3Q88UE1, HIS5_LACPLQ9RDX3, HIS5_LEGPN
Q6AE14, HIS5_LEIXXQ9ZGM1, HIS5_LEPBOP61780, HIS5_LEPIC
P59118, HIS5_LEPINQ9S4H8, HIS5_LEPIRQ92E86, HIS5_LISIN
C1L0J4, HIS5_LISMCQ722Y5, HIS5_LISMFB8DA60, HIS5_LISMH
Q8Y9G3, HIS5_LISMOA0AG17, HIS5_LISW6Q65RB8, HIS5_MANSM
C6C1X6, HIS5_MARSDQ8TS91, HIS5_METACQ465U3, HIS5_METBF
Q603K1, HIS5_METCAQ57929, HIS5_METJAQ8TV83, HIS5_METKA
Q8PVD5, HIS5_METMAO27568, HIS5_METTHB0JFQ9, HIS5_MICAN
Q2RGW0, HIS5_MOOTAP59957, HIS5_MYCBOQ9X7C0, HIS5_MYCLE
P60600, HIS5_MYCPAP9WMM0, HIS5_MYCTOP9WMM1, HIS5_MYCTU
Q3IUP8, HIS5_NATPDQ5FA21, HIS5_NEIG1Q9JVH3, HIS5_NEIMA
Q9K0H2, HIS5_NEIMBQ820D2, HIS5_NITEUQ2YAU8, HIS5_NITMU
Q3J6Q3, HIS5_NITOCQ5YYP7, HIS5_NOCFAB2J1A3, HIS5_NOSP7
Q8YX49, HIS5_NOSS1Q8ESS0, HIS5_OCEIHQ316L3, HIS5_OLEA2
P57921, HIS5_PASMUQ6D408, HIS5_PECASQ4FNT5, HIS5_PELUB
Q7N6I3, HIS5_PHOLLP62448, HIS5_PHOPRQ6KZD3, HIS5_PICTO
A9BB49, HIS5_PROM4Q31AG5, HIS5_PROM9Q7VBF5, HIS5_PROMA
Q7V127, HIS5_PROMPQ46JZ5, HIS5_PROMTQ48C78, HIS5_PSE14
Q4KJS7, HIS5_PSEF5Q3KJI8, HIS5_PSEPFQ88R44, HIS5_PSEPK
Q87UG1, HIS5_PSESMQ4ZLP9, HIS5_PSEU2Q4FPU1, HIS5_PSYA2
Q8ZY40, HIS5_PYRAEP58789, HIS5_PYRFUQ8XV83, HIS5_RALN1
A9WQ58, HIS5_RENSMQ98CT5, HIS5_RHILOQ92TB1, HIS5_RHIME
Q7ULP3, HIS5_RHOBAP60601, HIS5_RHOPAQ2RNA5, HIS5_RHORT
Q5LU93, HIS5_RUEPOO33777, HIS5_SACS2Q57MS0, HIS5_SALCH
Q5PDP6, HIS5_SALPAQ2S299, HIS5_SALRDP0A1R5, HIS5_SALTI
P0A1R4, HIS5_SALTYO94303, HIS5_SCHPOA3QF19, HIS5_SHELP
Q8EFB4, HIS5_SHEONA8FWC9, HIS5_SHESHQ323I9, HIS5_SHIBS
Q32EF2, HIS5_SHIDSQ83KJ5, HIS5_SHIFLQ3Z0G2, HIS5_SHISS
Q2YZ69, HIS5_STAABQ5HCM2, HIS5_STAACP64363, HIS5_STAAM
P64364, HIS5_STAANQ6GDC9, HIS5_STAARQ6G600, HIS5_STAAS
P64365, HIS5_STAAWQ5HKP0, HIS5_STAEQQ8CTV0, HIS5_STAES
Q4A047, HIS5_STAS1Q82AA2, HIS5_STRAWP16249, HIS5_STRCO
A8AY26, HIS5_STRGCQ8DTR1, HIS5_STRMUA3CNT2, HIS5_STRSV
Q4J8I5, HIS5_SULACQ30PY0, HIS5_SULDNQ970Y7, HIS5_SULTO
Q67KH8, HIS5_SYMTHQ3A135, HIS5_SYNC1Q31N58, HIS5_SYNE7
Q2JVR1, HIS5_SYNJAQ2JN09, HIS5_SYNJBQ5MZ63, HIS5_SYNP6
Q3AYZ0, HIS5_SYNS9Q3AI95, HIS5_SYNSCQ55805, HIS5_SYNY3
Q47QS5, HIS5_THEFYQ5JFV0, HIS5_THEKOQ9X0C8, HIS5_THEMA
P61781, HIS5_THET2Q7SIC0, HIS5_THET8Q8DIP5, HIS5_THEVB
Q3SEU6, HIS5_THIDAQ113E7, HIS5_TRIEIQ3M5W4, HIS5_TRIV2
Q9KSX0, HIS5_VIBCHQ87QK8, HIS5_VIBPAQ8D8Q3, HIS5_VIBVU
Q7M9X0, HIS5_WOLSUQ8PLG8, HIS5_XANACQ3BUF4, HIS5_XANC5
Q4UU43, HIS5_XANC8Q8P9P1, HIS5_XANCPQ2P3K0, HIS5_XANOM
Q5H0K8, HIS5_XANORQ9PBC8, HIS5_XYLFAQ87C32, HIS5_XYLFT
P33734, HIS5_YEASTQ8ZFX8, HIS5_YERPEQ66C52, HIS5_YERPS
Q5NMD4, HIS5_ZYMMOQ9PM72, HIS61_CAMJEQ7U5M8, HIS61_PARMW
Q7V8Q8, HIS61_PROMMQ9HU44, HIS61_PSEAEQ7MLS1, HIS61_VIBVY
Q0P8U3, HIS62_CAMJEA1W0U8, HIS62_CAMJJQ7U923, HIS62_PARMW
Q7V958, HIS62_PROMMP72139, HIS62_PSEAEQ7MPP3, HIS62_VIBVY
Q7MPK7, HIS63_VIBVYA3DJF7, HIS6_ACET2Q6F799, HIS6_ACIAD
B7GVJ5, HIS6_ACIB3B7IBD7, HIS6_ACIB5B2I0P3, HIS6_ACIBC
A3M9P1, HIS6_ACIBTC1FAD8, HIS6_ACIC5A5FYE5, HIS6_ACICJ
B9MDW3, HIS6_ACIETB7JA19, HIS6_ACIF2B5EQF2, HIS6_ACIF5
A1W444, HIS6_ACISJB3GZH3, HIS6_ACTP7A6VQ74, HIS6_ACTSZ
A0KKB0, HIS6_AERHHA4SMQ4, HIS6_AERS4B6JCG5, HIS6_AFIC5
P58799, HIS6_AGRFCB2UQA2, HIS6_AKKM8Q21U91, HIS6_ALBFT
Q0VM72, HIS6_ALCBSA8EQW5, HIS6_ALIB4Q5E633, HIS6_ALIF1
B5FDA4, HIS6_ALIFMB6EJ93, HIS6_ALISLQ5WDI2, HIS6_ALKCK
Q0A5D3, HIS6_ALKEHA6TKT6, HIS6_ALKMQB4RU62, HIS6_ALTMD
B7GL45, HIS6_ANOFWO66567, HIS6_AQUAEO29439, HIS6_ARCFU
Q5P795, HIS6_AROAEA0JVK5, HIS6_ARTS2P26721, HIS6_AZOBR
A8HYT7, HIS6_AZOC5B6YQ30, HIS6_AZOPCA1KAV4, HIS6_AZOSB
C1DJD3, HIS6_AZOVDC3P506, HIS6_BACAAC3L9P7, HIS6_BACAC
A0RBM2, HIS6_BACAHQ81T58, HIS6_BACANB7JFZ5, HIS6_BACC0
P62449, HIS6_BACC1B7INA3, HIS6_BACC2C1EMQ7, HIS6_BACC3
B7HHG5, HIS6_BACC4B7HKD4, HIS6_BACC7A7GMV0, HIS6_BACCN
B9IV00, HIS6_BACCQQ81G02, HIS6_BACCRQ63DW8, HIS6_BACCZ
Q5LBD7, HIS6_BACFNQ64RT2, HIS6_BACFRQ6HLE3, HIS6_BACHK
Q65EG3, HIS6_BACLDA9VLH7, HIS6_BACMKA8FHQ9, HIS6_BACP2
O34727, HIS6_BACSUQ8A7Z6, HIS6_BACTNA7Z962, HIS6_BACVZ
Q1LT72, HIS6_BAUCHB2ICL0, HIS6_BEII9C5BV99, HIS6_BEUC1
B8DUB2, HIS6_BIFA0A1A1I5, HIS6_BIFAAB3DRT8, HIS6_BIFLD
Q8G6F7, HIS6_BIFLOB7GQS5, HIS6_BIFLSQ7VQW5, HIS6_BLOFL
Q494D6, HIS6_BLOPBQ2KTT1, HIS6_BORA1Q7WDX9, HIS6_BORBR
Q7W2X9, HIS6_BORPAQ7VSY6, HIS6_BORPEQ89WM4, HIS6_BRADU
A5E8E5, HIS6_BRASBA4YJV5, HIS6_BRASOC0Z6P4, HIS6_BREBN
B2S984, HIS6_BRUA1Q2YQY8, HIS6_BRUA2A6WX52, HIS6_BRUA4
Q57AH3, HIS6_BRUABA9M9R9, HIS6_BRUC2C0RFX3, HIS6_BRUMB
Q8YE37, HIS6_BRUMEA5VT43, HIS6_BRUO2B0CJI6, HIS6_BRUSI
Q8FY07, HIS6_BRUSUP57206, HIS6_BUCAIQ9ZHE1, HIS6_BUCAP
P59521, HIS6_BUCBPB1YRW1, HIS6_BURA4A0K3V8, HIS6_BURCH
B4E641, HIS6_BURCJQ0BIW4, HIS6_BURCMQ39K85, HIS6_BURL3
Q845U7, HIS6_BURM1A3MPU4, HIS6_BURM7A2S754, HIS6_BURM9
Q62GE5, HIS6_BURMAA1V8H5, HIS6_BURMSB1JUA5, HIS6_BURO0
Q1BS31, HIS6_BURO1A3P027, HIS6_BURP0Q3JN02, HIS6_BURP1
A3NE93, HIS6_BURP6Q63Q92, HIS6_BURPSQ2SUA8, HIS6_BURTA
A4JAW9, HIS6_BURVGQ8R885, HIS6_CALS4A7ZGB2, HIS6_CAMC1
A7GVW5, HIS6_CAMC5A0RRT8, HIS6_CAMFFA7I416, HIS6_CAMHC
A8FNR4, HIS6_CAMJ8A7H5V4, HIS6_CAMJDQ5HSI8, HIS6_CAMJR
B9KEI4, HIS6_CAMLRQ3AD48, HIS6_CARHZB0T798, HIS6_CAUSK
Q9A229, HIS6_CAUVCB8GW15, HIS6_CAUVNB3PGA4, HIS6_CELJU
A3PI62, HIS6_CERS1P50937, HIS6_CERS4A4WUS8, HIS6_CERS5
B9KPC8, HIS6_CERSKQ11CK7, HIS6_CHESBA9WD80, HIS6_CHLAA
B8GBF3, HIS6_CHLADQ3ATG0, HIS6_CHLCHB3EEF2, HIS6_CHLL2
Q3B2Q6, HIS6_CHLL3B3QQ00, HIS6_CHLP8B3EKW7, HIS6_CHLPB
A1BHP6, HIS6_CHLPDA4SFU1, HIS6_CHLPMB9LFN2, HIS6_CHLSY
B3QSI0, HIS6_CHLT3Q8KCB0, HIS6_CHLTEA0M283, HIS6_CHRFK
Q1R080, HIS6_CHRSDQ7P0E9, HIS6_CHRVOB5EDR4, HIS6_CITBB
A8AEJ9, HIS6_CITK8A5CRW2, HIS6_CLAM3Q97KH8, HIS6_CLOAB
A7FU82, HIS6_CLOB1C3KVX6, HIS6_CLOB6A6LT22, HIS6_CLOB8
B2UX20, HIS6_CLOBAA5I246, HIS6_CLOBHC1FN42, HIS6_CLOBJ
B1ILB0, HIS6_CLOBKA7GDQ7, HIS6_CLOBLQ18C74, HIS6_CLOD6
Q47XB3, HIS6_COLP3C3PHA6, HIS6_CORA7P60714, HIS6_CORDI
Q8FNZ9, HIS6_COREFA4QFF9, HIS6_CORGBO31139, HIS6_CORGL
Q4JW52, HIS6_CORJKB1VH88, HIS6_CORU7A7MHE1, HIS6_CROS8
Q1LIB2, HIS6_CUPMCQ0K693, HIS6_CUPNHQ46WL8, HIS6_CUPPJ
B3R794, HIS6_CUPTRQ6A7Y8, HIS6_CUTAKQ11VM1, HIS6_CYTH3
Q47AM3, HIS6_DECARQ3Z7F5, HIS6_DEHM1A5FQI9, HIS6_DEHMB
Q3ZY29, HIS6_DEHMCQ1IX39, HIS6_DEIGDQ9RWD7, HIS6_DEIRA
A9BXA1, HIS6_DELASC0Q9D3, HIS6_DESAHB8FFL4, HIS6_DESAL
B1I556, HIS6_DESAPB8J216, HIS6_DESDAB8FP24, HIS6_DESHD
Q24QJ5, HIS6_DESHYA8ZVD2, HIS6_DESOHA4J707, HIS6_DESRM
P62450, HIS6_DESVHB8DQX6, HIS6_DESVMA1VGY9, HIS6_DESVV
B5YE90, HIS6_DICT6B8E2C8, HIS6_DICTDA8LHX5, HIS6_DINSH
A7ZNJ7, HIS6_ECO24B7UT62, HIS6_ECO27B7MDH9, HIS6_ECO45
B7L9Q2, HIS6_ECO55P60665, HIS6_ECO57B5YU81, HIS6_ECO5E
B7NQG5, HIS6_ECO7IB7MWU3, HIS6_ECO81B7M404, HIS6_ECO8A
C4ZSB4, HIS6_ECOBWB1X6W2, HIS6_ECODHA8A1P9, HIS6_ECOHS
A1ACN7, HIS6_ECOK1Q0TG62, HIS6_ECOL5Q8FG48, HIS6_ECOL6
B1IZ49, HIS6_ECOLCP60664, HIS6_ECOLIB7NC65, HIS6_ECOLU
B6I896, HIS6_ECOSEB1LP16, HIS6_ECOSMQ1RA48, HIS6_ECOUT
C5BG16, HIS6_EDWI9B2KEU8, HIS6_ELUMPB1H0E6, HIS6_ENDTX
A4WC74, HIS6_ENT38B7LUF6, HIS6_ESCF3B1YL09, HIS6_EXIS2
C4L176, HIS6_EXISAA5FFX6, HIS6_FLAJ1A6GY75, HIS6_FLAPJ
Q2J8L3, HIS6_FRACCB0TY41, HIS6_FRAP2B9M0M0, HIS6_GEODF
Q5KVD0, HIS6_GEOKAQ39YP2, HIS6_GEOMGP60715, HIS6_GEOSL
C6E7F4, HIS6_GEOSMC5D7N8, HIS6_GEOSWA4ISR2, HIS6_GEOTN
A5G8S9, HIS6_GEOURQ7NDC1, HIS6_GLOVIA9GZW9, HIS6_GLUDA
Q5FTN3, HIS6_GLUOXQ4QN69, HIS6_HAEI8A5UA15, HIS6_HAEIE
P44436, HIS6_HAEINQ2SMB2, HIS6_HAHCHQ9K6Z6, HIS6_HALH5
A1WW05, HIS6_HALHLQ5V2C9, HIS6_HALMAB8D116, HIS6_HALOH
B0R4B2, HIS6_HALS3Q9HR52, HIS6_HALSAQ18JI3, HIS6_HALWD
Q7VGZ1, HIS6_HELHPB0TDM7, HIS6_HELMIQ8GDQ6, HIS6_HELMO
A9B5I5, HIS6_HERA2A4G9I6, HIS6_HERARQ31E64, HIS6_HYDCU
B4U7M4, HIS6_HYDS0Q0C643, HIS6_HYPNAQ5QWQ5, HIS6_IDILO
A8AAK3, HIS6_IGNH4A6T376, HIS6_JANMAQ28NK0, HIS6_JANSC
A6WC96, HIS6_KINRDP45603, HIS6_KLEOXB5XPE2, HIS6_KLEP3
A6TBC8, HIS6_KLEP7B2GGU9, HIS6_KOCRDQ1IKB2, HIS6_KORVE
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B1JPW5, HIS6_YERPYQ5NMD6, HIS6_ZYMMO

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